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Susan E. HayhurstDepartment of Wildlife Health Sciences, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, 3001 Connecticut Ave NW, Washington, DC 20008.

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Jason D. EngelDepartment of Wildlife Health Sciences, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, 3001 Connecticut Ave NW, Washington, DC 20008.

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Timothy WalshDepartment of Wildlife Health Sciences, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, 3001 Connecticut Ave NW, Washington, DC 20008.

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Sebastian GordonDepartment of Wildlife Health Sciences, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, 3001 Connecticut Ave NW, Washington, DC 20008.

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Suzan MurrayDepartment of Wildlife Health Sciences, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, 3001 Connecticut Ave NW, Washington, DC 20008.

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Nancy BoedekerDepartment of Wildlife Health Sciences, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, 3001 Connecticut Ave NW, Washington, DC 20008.

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Contributor Notes

Dr. Engel's present address is Urologic Surgeons of Washington, 1147 20th St NW Ste 400, Washington, DC 20036. Dr. Hayhurst's present address is Mann Memorial Veterinary Clinic, PO Box 919, Kennebunk, ME 04043.

No external funding or financial support.

The authors thank Dr. Cathy Williams, Dr. Robert Schopler, and Dr. Matthew Nelson for sharing their expertise during the preparation of this report.

Address correspondence to Dr. Hayhurst (susan.hayhurst@gmail.com).